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In botany, a tree is a perennial woody plant with a single main trunk taller than about 2 metres, with xylem tissue in the trunk and branches that continues to enlarge during the life of the plant by the process of secondary growth.

In common usage, the definition of tree may be narrower, including only very tall plants or those that could yield significant wood. The definition can instead be broader, including tall plants without secondary growth such as the taller palms and the tree ferns. Bamboo is a tall woody plant that is not usually considered to be a tree.

A tree typically has many secondary branches supported clear of the ground by the trunk. This trunk typically contains woody tissue for strength, and vascular tissue to carry materials from one part of the tree to another. For most trees it is surrounded by a layer of bark which serves as a protective barrier. Below the ground, the roots branch and spread out widely; they serve to anchor the tree and extract moisture and nutrients from the soil. Above ground, the branches divide into smaller branches and shoots. The shoots typically bear leaves, which capture light energy and convert it into chemical energy by photosynthesis, providing the food needed by the tree for its growth and development.

Flowers and fruit may also be present, but some trees such as conifers instead have pollen cones and seed cones, and others such as tree ferns produce spores instead.

Trees tend to be long-lived, some reaching several thousand years old. The tallest known specimen on Earth is 115.6 m (379 ft) and they have a theoretical maximum height of 130m (426 ft). Trees have been in existence on the Earth for 370 million years. Trees are not a taxonomic group but are a number of plant species that have independently evolved a woody trunk and branches as a way to tower above other plants and make full use of the sunlight.

Trees play a significant part in reducing erosion and moderating the climate. They remove carbon dioxide from the atmosphere and store large quantities of carbon in their tissues. Trees and forests provide a habitat for many species of animals and plants. Tropical rainforests are one of the most biodiverse habitats in the world. Trees provide shade and shelter, timber for construction, fuel for cooking and heating, and fruit for food as well as having many other uses. In parts of the world, forests are shrinking as trees are cleared to increase the amount of land available for agriculture. Because of their longevity and usefulness, trees have always been revered and they play a role in many of the world's mythologies.